Cucujiformia is an infraorder of polyphagan beetles, representing most plant-eating beetles. It includes species from the major superfamilies of Chrysomeloidea, Curculionoidea, Cucujoidea, and Tenebrionoidea.
Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles (Scolytinae), which are weevils.
Corylophidae is a family of minute hooded beetles, sometimes called minute fungus beetles, in the superfamily Coccinelloidea. There are about 18 genera and at least 120 described species in Corylophidae. They feed on microfungi such as molds, and are often found associated with bark, as well as in leaf litter and other decaying vegetation. In older literature, the family name was often given as Orthoperidae.
Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.
Boganiidae is a family of beetles, in the superfamily Cucujoidea. Members of the family are found in southern Africa, Australia and New Caledonia. Adults and larvae are pollenivorous, feeding on the pollen of cycads and flowering plants of the families Myrtaceae, Meliaceae, Cunoniaceae and Elaeocarpaceae. Metacucujus and Paracucujus act as pollinators for cycads Encephalartos and Macrozamia respectively. This association with cycads goes back to at least the Mid-Cretaceous, with an extinct form being found with preserved cycad pollen in 99 million year old Burmese amber.
Bothrideridae is a family of beetles in the superfamily Coccinelloidea. They are known commonly as the cocoon-forming beetles or dry bark beetles. They occur worldwide with most native to the Old World tropics. In older literature, the family was often included in the family Colydiidae, but is now considered unrelated.
Hobartiidae is a family of beetles, in the superfamily Cucujoidea. There are only two known genera Hobartius and Hydnobioides, with six species, five of which are native to Australia, and one species of Hobartius native to Chile and Argentina in South America. Members of the family are mycophagous, living in rotten, fungus infested logs of Araucaria, Nothofagus, and Eucalyptus, where they feed on the fruiting bodies of basidiomycetes.
Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, Helota MacLeay, Neohelota Ohta, Afrohelotina Kirejtshuk, Metahelotella Kirejtshuk, and Strophohelota Kirejtshuk. Helotidae are found mainly in the Old World tropics and are absent from Australia and Madagascar. The antennae are clubbed on the final three segments and is retractable within grooves under the head. The wings have reduced venation with just 4 anal veins. Helotids are known to be associated with sap, fruit and flowers, and the larvae of some species are known to bore into wood in order to pupate.
Protocucujidae is a family of beetles, in the superfamily Cucujoidea. It has a single known genus, Ericmodes. Species of Ericmodes are native to southern South America and Eastern Australia. Little is known of their biology, though adults and larvae probably live on vegetation, with adults having also been found in leaf litter and in flight.
Anamorphidae is a family of beetles in the superfamily Coccinelloidea, formerly included within the family Endomychidae. They are found worldwide. Like enchomyids, they are fungivores, with adult and larval stages thought to exclusively consume fungal spores.
Eupsilobiidae is a family of beetles in the superfamily Coccinelloidea, formerly included within the family Endomychidae. Most genera are restricted to the Neotropics, while the genus Eidoreus is found worldwide. They are fungivores, and have been observed living commensally in bee and ant nests.
Bystus is a genus of beetles in the family Anamorphidae. There are about six described species in Bystus.
Parmulini is a tribe of minute hooded beetles in the family Corylophidae. There are at least 2 genera and more than 40 described species in Parmulini.
Arthrolips is a genus of minute hooded beetles in the family Corylophidae. There are more than 20 described species in Arthrolips.
Sericoderus is a genus of minute hooded beetles in the family Corylophidae. There are about 12 described species in Sericoderus.
Aenigmaticum is a genus of minute hooded beetles in the family Corylophidae. There are about five described species in Aenigmaticum.
Rypobius is a genus of minute hooded beetles in the family Corylophidae. There are about five described species in Rypobius.
Rypobiini is a tribe of minute hooded beetles in the family Corylophidae. There are at least 2 genera and about 14 described species in Rypobiini.
Merophysiinae is a subfamily of handsome fungus beetles in the family Endomychidae.
Myrabolia is the only genus in the beetle family Myraboliidae in the superfamily Cucujoidea. It has about 13 species, found in Australia. Adults and possibly larvae live under the bark of Eucalyptus trees.
