| Leiosauridae | |
|---|---|
 | |
| Diplolaemus bibronii | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | Squamata |
| Suborder: | Iguania |
| Clade: | Pleurodonta |
| Family: | Leiosauridae Frost, Etheridge, Janies, & Titus, 2001 |
Leiosauridae is a family of iguanian lizards containing six genera and 34 species. The family is endemic to Central America and South America.
A 2022 phylogenetic study found the Leiosauridae to be the sister taxa to the Opluridae, a unique family of iguanians endemic to Madagascar, and one of only two pleurodont lineages found outside the Americas (the other being the genus Brachylophus ). This called into question the previous hypothesis of Opluridae being the last members of an ancient lineage of Gondwanan iguanians, as the study found that the Leiosauridae and Opluridae only diverged during the Paleocene, about 60 million years ago. Opluridae are now thought to have colonized Madagascar via oceanic dispersal from South America, either directly or indirectly. [1]
The family Leiosauridae contains the following genera: [2]
| Genus | Image | Type species | Taxon author | Common name | Species |
|---|---|---|---|---|---|
| Anisolepis | A. undulatus (Wiegmann, 1834) | Boulenger, 1885 | Tree lizards | 3 | |
| Diplolaemus |  D. darwinii | D. darwinii Bell, 1843 | Bell, 1843 | Patagonian lizards | 4 |
| Enyalius |  E. catenatus | E. catenatus (Wied, 1821) | Wagler, 1830 | Fathead anoles | 11 |
| Leiosaurus |  L. bellii | L. bellii Duméril & Bibron, 1837 | Duméril & Bibron, 1837 | Smooth iguanas | 4 |
| Pristidactylus |  P. achalensis | P. fasciatus (d'Orbigny & Bibron, 1837) | Fitzinger, 1843 | Big-headed iguanas | 10 |
| Urostrophus | U. vautieri Duméril & Bibron, 1837 | Duméril & Bibron, 1837 | Steppe iguanas | 2 |
Squamata is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians, which are collectively known as squamates or scaled reptiles. With over 11,500 species, it is also the second-largest order of extant (living) vertebrates, after the perciform fish. Members of the order are distinguished by their skins, which bear horny scales or shields, and must periodically engage in molting. They also possess movable quadrate bones, making possible movement of the upper jaw relative to the neurocranium. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey. Squamates are the most variably sized living reptiles, ranging from the 16 mm (0.63 in) dwarf gecko to the 6.5 m (21 ft) reticulated python. The now-extinct mosasaurs reached lengths over 14 m (46 ft).
Gekkonidae is the largest family of geckos, containing over 950 described species in 64 genera. The Gekkonidae contain many of the most widespread gecko species, including house geckos (Hemidactylus), the tokay gecko (Gekko), day geckos (Phelsuma), the mourning gecko (Lepidodactylus), and dtellas (Gehyra). Gekkonid geckos occur globally and are particularly diverse in tropical areas.
Agamidae is a family of over 550 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards.
Corytophanidae is a family of iguanian lizards, also called casquehead lizards or helmeted lizards, endemic to the New World. Nine species of casquehead lizards from three genera are recognized.
The Crotaphytidae, or collared lizards, are a family of desert-dwelling reptiles native to the Southwestern United States and northern Mexico. Alternatively they are recognized as a subfamily, Crotaphytinae, within the clade Pleurodonta. They are very fast-moving animals, with long limbs and tails; some species are capable of achieving bipedal running at top speed. This species is carnivorous, feeding mainly on insects and smaller lizards. The two genera contain 12 species.
The Opluridae, or Madagascan iguanas, are a family of moderately sized lizards native to Madagascar and Grande Comore. There are eight species in two genera, with most of the species being in Oplurus. The Opluridae, along with Brachylophus of Fiji, are the only extant members of the Pleurodonta that are found outside the Americas.
The Iguanidae is a family of lizards composed of the iguanas, chuckwallas, and their prehistoric relatives, including the widespread green iguana.
The genus Brachylophus consists of four extant iguanid species native to the islands of Fiji and a giant extinct species from Tonga in the South West Pacific. One of the extant species, B. fasciatus, is also present on Tonga, where it has apparently been introduced by humans.
Iguania is an infraorder of squamate reptiles that includes iguanas, chameleons, agamids, and New World lizards like anoles and phrynosomatids. Using morphological features as a guide to evolutionary relationships, the Iguania are believed to form the sister group to the remainder of the Squamata, which comprise nearly 11,000 named species, roughly 2000 of which are iguanians. However, molecular information has placed Iguania well within the Squamata as sister taxa to the Anguimorpha and closely related to snakes. The order has been under debate and revisions after being classified by Charles Lewis Camp in 1923 due to difficulties finding adequate synapomorphic morphological characteristics. Most Iguanias are arboreal but there are several terrestrial groups. They usually have primitive fleshy, non-prehensile tongues, although the tongue is highly modified in chameleons. The group has a fossil record that extends back to the Early Jurassic. Today they are scattered occurring in Madagascar, the Fiji and Friendly Islands and Western Hemisphere.
In phylogenetics, basal is the direction of the base of a rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa.
The Madagascar girdled lizard or Madagascar plated lizard is a species of lizard in the family Gerrhosauridae. It is found in Madagascar, some outlying islands and the Seychelles (Cosmoledo). This generally common lizard is found in a range of habitat types, including open and degraded humid forest at mid and low altitudes. It also occurs in plantations. On Cosmoledo, it occurs in open shrubland. This oviparous lizard grows to 30 cm (12 in) total length.
Diploglossidae is a family of anguimorph lizards native to the Americas, with most genera being endemic to Hispaniola. Most members of this family are known as galliwasps. They were formerly considered a subfamily of Anguidae, but genetic evidence has shown them to be less closely related to other members of Anguidae than Anniellidae is.
Iguana is a genus of herbivorous lizards that are native to tropical areas of Mexico, Central America, South America, and the Caribbean. The genus was first described in 1768 by Austrian naturalist Josephus Nicolaus Laurenti in his book Specimen Medicum, Exhibens Synopsin Reptilium Emendatam cum Experimentis circa Venena. Three species are placed in the genus: the green iguana, which is widespread throughout its range and a popular pet, the marine iguana and the Lesser Antillean iguana, which is native to the Lesser Antilles. Genetic analysis indicates that the green iguana may comprise a complex of multiple species, some of which have been recently described, but the Reptile Database considers all of these as subspecies of the green iguana.
Scleroglossa is a group of lizards that includes geckos, autarchoglossans, and amphisbaenians. Scleroglossa is supported by phylogenetic analyses that use morphological features. According to most morphological analyses, Scleroglossa is the sister group of the clade Iguania, which includes iguanas, chameleons, agamids, and New World lizards. Together, Scleroglossa and Iguania make up the crown group Squamata, the smallest evolutionary grouping to include all living lizards and snakes.
The Diplodactylidae are a family in the suborder Gekkota (geckos), with over 150 species in 25 genera. These geckos occur in Australia, New Zealand, and New Caledonia. Diplodactylids are the most ecologically diverse and widespread family of geckos in both Australia and New Caledonia, and are the only family of geckos found in New Zealand. Three diplodactylid genera have recently been split into multiple new genera.
Polyglyphanodontia, also known as the Borioteiioidea, is an extinct clade of lizards from the Cretaceous that includes around a dozen genera. Polyglyphanodontians were the dominant group of lizards in North America and Asia during the Late Cretaceous. Most polyglyphanodontians are Late Cretaceous in age, though the oldest one, Kuwajimalla kagaensis, is known from the Early Cretaceous Kuwajima Formation (Japan). Early Cretaceous South American taxon Tijubina, and possibly also Olindalacerta, might also fall within Polyglyphanodontia or be closely allied to the group, but if so, they would be two of only four Gondwanan examples of an otherwise Laurasian clade. They produced a remarkable range of forms. Chamopsiids, including Chamops, were characterized by large, blunt, crushing teeth, and were most likely omnivores. Polyglyphanodon, from the Maastrichtian of Utah, was another herbivore, but its teeth formed a series of transverse blades, similar to those of Trilophosaurus. Peneteius had remarkable, multicusped teeth, similar to those of mammals. The polyglyphanodontids first appear in the latter part of the Early Cretaceous in North America, and became extinct during the Cretaceous-Paleogene extinction event. Polyglyphanodontians closely resembled the teiid lizards, and purported teiid lizards from the Late Cretaceous appear to be polyglyphanodontians. The only species known to have survived the Cretaceous was Chamops, which survived until the very early Ypresian.
Gobiguania is an extinct clade of iguanian lizards from the Late Cretaceous. All known gobiguanians are endemic to the Gobi Desert of Mongolia. Gobiguania was given a phylogenetic definition by Jack Conrad and Mark Norell in 2007 as all taxa more closely related to Anchaurosaurus gilmorei than to Iguana iguana, Crotaphytus collaris, or Agama agama. According to Conrad and Norell's phylogenetic analysis, Gobiguania includes Anchaurosaurus as well as several other Late Cretaceous lizards such as Ctenomastax, Temujinia, Saichangurvel, and Zapsosaurus. A phylogenetic analysis published in 2012 indicated that three other lizard genera — Igua, Isodontosaurus, and Polrussia, all from Mongolia and all from the Late Cretaceous — are also gobiguanians. Below is a cladogram from the analysis:
Priscagamidae is an extinct family of iguanian lizards known from the Late Cretaceous of Mongolia and China and the Eocene of India, spanning a range from 83.6 to 48.6 million years ago. Probably the earliest priscagamids on indeterminate genera were found in Aptian-Albian sediments in "Hobur", Mongolia. It includes the genera Heterodontagama, Mimeosaurus, Phrynosomimus, Priscagama, and possibly Pleurodontagama. The first fossils of priscagamids were found in the Djadochta and Khermeen Tsav formations of Mongolia. More recently they have been found in the Cambay Formation in India, leading to the naming of Heterodontagama in 2013. Priscagamidae was originally described as a subfamily of Agamidae called Priscagaminae in 1984, but it was reclassified as a distinct family in 1989. Most phylogenetic analyses still find a close relationship between Priscagamidae and Agamidae, although a 2015 study found it to be basal to all other iguanian clades, warranting its removal from Iguania and placement in a larger clade called Iguanomorpha.
Acrodonta are a subclade of iguanian squamates consisting almost entirely of Old World taxa. Extant representation include the families Chamaeleonidae (chameleons) and Agamidae, with at least over 500 species described. A fossil genus, Gueragama, was found in Brazil, making it the only known American representative of the group.
Enyalius boulengeri, also known commonly as the Brazilian fathead anole, is a species of lizard in the family Leiosauridae. The species is endemic to Brazil.
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