Lemmysuchus

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Lemmysuchus
Temporal range: Late Jurassic, Callovian
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Lemmysuchus obtusidens.png
Holotype skull in multiple views
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pseudosuchia
Superorder: Crocodylomorpha
Suborder: Thalattosuchia
Family: Machimosauridae
Subfamily: Machimosaurinae
Tribe: Machimosaurini
Genus: Lemmysuchus
Johnson, Young et al., 2017
Species:
L. obtusidens
Binomial name
Lemmysuchus obtusidens
Andrews, 1909
Synonyms

Lemmysuchus is a genus of machimosaurid thalattosuchian from the Middle Jurassic Callovian of England and France. Like many other teleosauroids from Europe, it has had a convoluted taxonomic history.

History

Skull and other elements of the holotype specimen A descriptive catalogue of the marine reptiles of the Oxford clay. Based on the Leeds Collection in the British Museum (Natural History), London (1910) (20252762483).jpg
Skull and other elements of the holotype specimen

"Steneosaurus" obtusidens was first described from the Callovian-aged Oxford Clay Formation of Cambridgeshire, southeastern England on the basis of a partial skeleton (NHMUK R.3168). [1] Characters cited as diagnostic for "S." obtusidens included: short rostrum (preorbital length 52 percent of total skull length); teeth blunt and rounded at the tips; armor adorned with elongate pit ornamentation.

The validity of "Steneosaurus" obtusidens was disputed in a 1987 paper reviewing the type specimens of nominal teleosauroid species from the Oxford Clay. Characters used by Charles William Andrews to distinguish "S." obtusidens from other species were dismissed as variable within specimens of Steneosaurus edwardsi , and "S." obtusidens was sunk as a junior synonym of Steneosaurus durobrivensis (=S. edwardsi). [2] Some authors considered this taxon a probable junior synonym of Machimosaurus hughii based on a subsequently discovered specimen found in Callovian-age exposures in Calvados, Lower Normandy, France, though they stressed that a taxonomic review of blunt-snouted teleosaurids was needed. [3] [4] This synonymy was accepted in a 2009 paper regarding thalattosuchian morphometrics without comment. [5]

In 2013, a new specimen of Machimosaurus was described from Late Jurassic (Kimmeridgian deposits in southern Germany (later named Machimosaurus buffetauti), and it became clear that variation within blunt-toothed teleosauroids was taxonomically significant enough for "Steneosaurus" obtusidens to be recognized as generically distinct from Machimosaurus. [6] Other cladistic and comparative studies agreed with this assessment, recovering "S." obtusidens in a clade with Machimosaurus and Steneosaurus edwardsi. [7] [8] [9] [10]

In 2017, the species was moved to its own genus, Lemmysuchus, referring to Ian Fraser Kilmister, better known as "Lemmy", of the band Motörhead, [11] and the Greek word for crocodile, soukhos. [12]

Related Research Articles

<i>Dakosaurus</i> Metriorhynchid reptile genus from the Late Jurassic / Early Cretaceous period

Dakosaurus is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially. The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named Geosaurus maximus by Theodor Plieninger in 1846. Dakosaurus was a carnivore that spent much, if not all, its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests have been discovered that have been referred to Dakosaurus, whether it gave birth to live young at sea like dolphins and ichthyosaurs or came ashore like turtles is not known. The name Dakosaurus means "biter lizard", and is derived from the Greek dakos ("biter") and σαῦρος -sauros ("lizard").

<i>Machimosaurus</i> Genus of reptiles

Machimosaurus is an extinct genus of machimosaurid crocodyliform from the Late Jurassic and Early Cretaceous. The type species, Machimosaurus hugii, was found in Switzerland. Other fossils have been found in England, France, Germany, Portugal, Switzerland and Tunisia. Machimosaurus rex is the largest named teleosauroid and thalattosuchian, with an estimated length of approximately 7.2 metres. Machimosaurus is the largest known crocodyliform of the Jurassic.

<i>Metriorhynchus</i> Extinct genus of reptiles

Metriorhynchus is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, M. brevirostris was named in 1829 as a species of Steneosaurus before being named as a separate genus by the German palaeontologist Christian von Meyer in 1832. The name Metriorhynchus means "Moderate snout", and is derived from the Greek Metrio- ("moderate") and -rhynchos ("snout").

Metriorhynchidae Extinct family of reptiles

Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs.

Thalattosuchia Clade of marine crocodylomorphs

Thalattosuchia is the name given to a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin.

Teleosauridae Extinct family of reptiles

Teleosauridae is a family of extinct typically marine crocodylomorphs similar to the modern gharial that lived during the Jurassic period. Teleosaurids were thalattosuchians closely related to the fully aquatic metriorhynchoids, but were less adapted to an open-ocean, pelagic lifestyle. The family was originally coined to include all the semi-aquatic thalattosuchians and was equivalent to the modern superfamily Teleosauroidea. However, as teleosauroid relationships and diversity was better studied in the 21st century, the division of teleosauroids into two distinct evolutionary lineages led to the establishment of Teleosauridae as a more restrictive family within the group, together with its sister family Machimosauridae.

<i>Plesiosuchus</i> Extinct genus of reptiles

Plesiosuchus is an extinct genus of geosaurine metriorhynchid crocodyliform known from the Late Jurassic of Dorset, England and possibly also Spain. It contains a single species, Plesiosuchus manselii.

<i>Pelagosaurus</i> Extinct genus of reptiles

Pelagosaurus is an extinct genus of thalattosuchian crocodyliform that lived during the Toarcian stage of the Lower Jurassic, around 183 Ma to 176 Ma, in shallow epicontinental seas that covered much of what is now Western Europe. The systematic taxonomy of Pelagosaurus has been fiercely disputed over the years, and was assigned to Thalattosuchia after its systematics within Teleosauridae were disputed. Pelagosaurus measured 3 m (10 ft) in length with a weight of 450 kg (1000 lbs), and was markedly similar to the modern-day gharial, which has similar adaptions and carnivorous feeding habits.

Cricosaurus is an extinct genus of marine crocodyliforms of the Late Jurassic. belonging to the family Metriorhynchidae. The genus was established by Johann Andreas Wagner in 1858 for three skulls from the Tithonian of Germany. The name Cricosaurus means "Ring lizard", and is derived from the Greek Cricos- ("ring") and σαῦρος -sauros ("lizard").

<i>Steneosaurus</i> Extinct genus of reptiles

Steneosaurus is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic of France. The genus has been used as a wastebasket taxon for thalattosuchian fossils for over two centuries, and almost all known historical species of teleosauroid have been included within it at one point. The genus has remained a wastebasket, with numerous species still included under the label ‘Steneosaurus’, many of which are unrelated to each other.

<i>Mycterosuchus</i> Extinct genus of reptiles

Mycterosuchus is an extinct genus of teleosaurid crocodyliform from the Middle Jurassic (Callovian) of England. Although previously synonymized with Steneosaurus, recent cladistic analysis considers it distantly related to the Steneosaurus type species.

<i>Mystriosaurus</i> Extinct genus of crocodilians

Mystriosaurus is an extinct genus of teleosaurid crocodyliform from the Early Jurassic (Toarcian). Fossil specimens have been found in the Whitby Mudstone of England and Posidonia Shale of Germany. The only known species, M. laurillardi, exceeded 4 metres (13 ft) in length.

<i>Sericodon</i> Extinct genus of reptiles

Sericodon is an extinct genus of teleosaurid crocodyliform from the Late Jurassic (Tithonian) of Germany and Switzerland. The genus contains a single species, S. jugleri.

The Reuchenette Formation is a Jurassic geologic formation in Switzerland. It is Kimmeridgian in age and predominantly consists of well stratified limestone, with lithology variable both laterally and stratigraphically including wackestones, packstones and grainstones, as well as mudstone. Dinosaur remains are among the fossils that have been recovered from the formation, including the Turiasaurian sauropod Amanzia greppini, alongside a theropod tooth belonging to Ceratosauria indet, originally assigned to Megalosaurus meriani. teleosaurid crocodyliformes are also known, including Sericodon, Proexochokefalos and Machimosaurus. The thalassochelydian turtle Solnhofia is known from the formation, as is the platychelyid turtle Platychelys.

The Tereñes Formation or Tereñes Marl is a Late Jurassic (Kimmeridgian) geologic formation in Asturias, Spain. The grey marls of the formation were deposited in a lagoonal environment at a muddy coast along a temporary inland sea. The lower section of the formation comprises silty and chalky sandstones with desiccation cracks and ripple marks, then becomes a bituminous, prominently ostracod-bearing, pelecypod shell chalk, lime chalk marl and marl. Fossil tracks have been reported from the formation.

<i>Torvoneustes</i> Extinct genus of reptiles

Torvoneustes is an extinct genus of metriorhynchid thalattosuchian. It is known from skull and postcranial remains found in the Kimmeridge Clay Formation of Dorset and Wiltshire, England, and also from Oaxaca, Mexico . The holotype skull of the type species was initially assigned to the species Metriorhynchus superciliosus. Postcranial remains were later discovered from the same quarry as the skull, and then these specimens were recognised as belonging to a new species of Dakosaurus, as D. carpenteri. The species was named to honour Simon Carpenter, an amateur geologist from Frome in Somerset, who discovered the fossils.

<i>Macrospondylus</i> Extinct genus of reptiles

Macrospondylus is an extinct genus of machimosaurid teleosauroid crocodyliform from the Early Jurassic (Toarcian) of Europe. Fossils are known from the Posidonia Shale of Germany, the Whitby Mudstone of the United Kingdom, and the "schistes bitumineux" of Luxembourg.

<i>Bathysuchus</i> Extinct genus of reptiles

Bathysuchus is an extinct genus of teleosaurid thalattosuchian from Late Jurassic (Kimmeridgian) deep water marine deposits in England and France. Bathysuchus displays features that suggest it was more pelagic than other teleosaurids, including smoother skull bones and reduced armour plating, similar to the fully marine metriorhynchids. This was possibly an adaptation to rising sea levels during the Kimmeridgian, as its earlier relatives such as Teleosaurus were suited for shallow coasts and lagoon environments.

<i>Charitomenosuchus</i> Extinct genus of reptiles

Charitomenosuchus is an extinct genus of machimosaurid teleosauroid from the Callovian Oxford Clay Formation of England.

<i>Proexochokefalos</i> Extinct genus of reptiles

Proexochokefalos is an extinct genus of machimosaurid teleosauroid from the Jurassic of France

References

  1. Andrews, C.W. (1909). "On some new Steneosaurs from the Oxford Clay of Peterborough". Annals and Magazine of Natural History. 3 (15): 299–308. doi:10.1080/00222930908692579.
  2. Adams-Tresman, S. M. (1987). "The Callovian (Middle Jurassic) teleosaurid marine crocodiles from central England". Palaeontology. 30 (1): 195–206.
  3. Hua, Vignaud; Pennetier; Pennetier (1994). "Un squelette de Steneosaurus obtusidens Andrews, 1909 dans le Callovien de Villers-sur-mer(Calvados, France)et le problème de la definition des Teleosauridae à dents obtuses". Comptes Rendus de l'Académie des Sciences, Série II. 318: 1557–1562.
  4. Hua, S., 1999, Le Crocodilien Machimosaurus mosae (Thalattosuchia, Teleosauridae) du Kimmeridgien du Boulonnais (Pas de Calais, France): Palaeontographica Abt. A., v. 252, Lfg. 4-6, p. 141-170.
  5. Pierce, S. E.; Angielczyk, K. D.; Rayfield, E. J. (2009). "Morphospace occupation in thalattosuchian crocodylomorphs: skull shape variation, species delineation and temporal patterns" (PDF). Palaeontology. 52 (5): 1057–1097. doi: 10.1111/j.1475-4983.2009.00904.x .
  6. Martin, J. E.; Vincent, P. (2013). "New remains of Machimosaurus hugii von Meyer, 1837 (Crocodilia, Thalattosuchia) from the Kimmeridgian of Germany". Fossil Record. 16 (2): 179–196. doi: 10.1002/mmng.201300009 .
  7. Mueller-Töwe, IJ (2005). "Phylogenetic relationships of the Thalattosuchia". Zitteliana. 45: 211–213.
  8. Young, M. T.; Brusatte, S. L.; De Andrade, M. B.; Desojo, J. B.; Beatty, B. L.; Steel, L.; Fernández, M. S.; Sakamoto, M.; Ruiz-Omeñaca, J. I.; Schoch, R. R. (2012). "Butler, Richard J, ed. "The Cranial Osteology and Feeding Ecology of the Metriorhynchid Crocodylomorph Genera Dakosaurus and Plesiosuchus from the Late Jurassic of Europe". PLOS ONE. 7 (9): e44985. doi: 10.1371/journal.pone.0044985 . PMC   3445579 . PMID   23028723.
  9. Young, M. T.; Hua, S.; Steel, L.; Foffa, D.; Brusatte, S. L.; Thüring, S.; Mateus, O.; Ignacio-Ruiz Omeñaca, J.; Lepage, Y.; Havilk, P.; Andrade, M. B. (2014). "Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)". Royal Society Open Science. 1 (140222): 1–42. Bibcode:2014RSOS....140222Y. doi:10.1098/rsos.140222. PMC   4448888 . PMID   26064545.
  10. Johnson, MM; Young, MT; Steel, L; Foffa, D; Smith, AS; Hua, S; Havlik, P; Howlett, EA; Dyke, G (2018). "Re-description of 'Steneosaurus' obustidens Andrews, 1909, an unusual macrophagous teleosaurid crocodylomorph from the Middle Jurassic of England". Zoological Journal of the Linnean Society. 182 (2): 385–418. doi:10.1093/zoolinnean/zlx035.
  11. "Scaly snuggle-toothed beast? That'll be Lemmy" > Daily Telegraph Issue no 50,455 dated Wednesday 9 August 2017
  12. Johnson, Michela M.; Young, Mark T.; Steel, Lorna; Foffa, Davide; Smith, Adam S.; Hua, StÉphane; Havlik, Philipe; Howlett, Eliza A.; Dyke, Gareth (2017). "Re-description of 'Steneosaurus' obtusidens Andrews, 1909, an unusual macrophagous teleosaurid crocodylomorph from the Middle Jurassic of England". Zoological Journal of the Linnean Society. 182 (2): 385–418. doi:10.1093/zoolinnean/zlx035.