Pinus radiata

Last updated

Monterey pine
Pinus radiata BigSur.jpg
Status TNC G1.svg
Critically Imperiled  (NatureServe) [2]
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Gymnospermae
Division: Pinophyta
Class: Pinopsida
Order: Pinales
Family: Pinaceae
Genus: Pinus
Subgenus: P. subg. Pinus
Section: P. sect. Trifoliae
Subsection: P. subsect. Australes
Species:
P. radiata
Binomial name
Pinus radiata
Pinus radiata range map 2.png
Natural range of Pinus radiata

Pinus radiata (syn. Pinus insignis), the Monterey pine, [3] insignis pine [4] or radiata pine, is a species of pine native to the Central Coast of California and Mexico (on Guadalupe Island and Cedros island). It is an evergreen conifer in the family Pinaceae.

Contents

P. radiata is a versatile, fast-growing, medium-density softwood, suitable for a wide range of uses and valued for rapid growth, as well as desirable lumber and pulp qualities. [5] Its silviculture reflects a century of research, observation and practice. [5] It is often considered a model for growers of other plantation species. [5]

Although P. radiata is extensively cultivated as a plantation timber in many temperate parts of the world, [6] it faces serious threats in its natural range, [7] due to the introduction of a fungal parasite, the pine pitch canker ( Fusarium circinatum ).

Description

Ovulate cone Pinus radiata cone.jpg
Ovulate cone
Pollen cones, 2 cm scale bar Pinus radiata pollen cones, 2 cm scale bar.png
Pollen cones, 2 cm scale bar

P. radiata is a coniferous evergreen tree growing to 15–30 m (50–100 ft) tall in the wild, but up to 60 m (200 ft) in cultivation in optimum conditions, with upward pointing branches and a rounded top. The leaves ("needles") are bright green, in clusters of three (two in var. binata), slender, 8–15 cm (3–6 in) long and with a blunt tip. The ovulate cones are 7–17 cm (3–6+12 in) long, brown, ovoid (egg-shaped), and usually set asymmetrically on a branch, attached at an oblique angle. The bark is fissured and dark grey to brown. When not cut short by disease or harvesting, it has a lifespan of 80 to 90 years.[ citation needed ]

The specific epithet radiata refers to the cracks which radiate from the umbo of the cone scales. [8]

It is closely related to bishop pine and knobcone pine, hybridizing readily with both species; it is distinguished from the former by needles in threes (not pairs), and from both by the cones not having a sharp spine on the scales.[ citation needed ]

The modern tree is vastly different from the native tree of Monterey. [8] [ self-published source? ] In plantations the tree is commonly planted at 4 square meter spacing on a wide variety of landscapes from flat to moderately steep hills. [8] Because of selective breeding and more recently the extensive use of growth factor seedlings, forests planted since the 1990s have very straight tall trunks without the problem of twin leaders. [8] The trees are pruned in three lifts so that the lower two-thirds of a mature tree is free of branches and hence of knots. [8]

Distribution and habitat

In the United States, it is native to three very limited areas located in Santa Cruz, Monterey, and San Luis Obispo Counties of California. [9]

In Mexico, it is found on two islands in the Pacific Ocean, Guadalupe Island and Cedros Island. [9] On Guadalupe Island, located 280 km (150 nmi) off of the mainland coast, the pines are found on the steep northern end of the island, at elevations of around 500 to 1,200 m (1,600 to 3,900 ft), where they follow the ridgetops and steep slopes. On Cedros Island, the pines are more abundant, being found in far greater numbers partly due to the lack of feral goats. They are found at lower elevations than on Guadalupe, at around 285 to 690 m (935 to 2,264 ft), on the windward ridges and canyons of the north and central parts of the island. In both cases, the pines seem to be heavily dependent on locations with a high frequency of fog. [10]

In Australia, New Zealand, and Spain it is the leading introduced tree [11] and in Argentina, Chile, Uruguay, Kenya, and South Africa it is a major plantation species. It is also an introduced tree on the world's most remote inhabited island, Tristan da Cunha.[ citation needed ]

Taxonomy

Subdivisions

Two varieties of this species have been recognized, each corresponding to the island they are endemic to in Baja California. Some authors lump these taxa and do not recognize infraspecies. [12]

On both islands it is the only pine species, and one of the few tree species. Compared to the mainland species, which are mostly three-needled, the insular varieties have their needles in bundles of two. Their cones are also smaller, and they show greater wind resistance in regards to wind-induced toppling. [10]

Ecology

Pinus radiata forest in Point Lobos, California Pinus radiata Lobos.jpg
Pinus radiata forest in Point Lobos, California

P. radiata is adapted to cope with stand-killing fire disturbance. Its cones are serotinous, i.e. they remain closed until opened by the heat of a forest fire; the abundant seeds are then discharged to regenerate on the burned forest floor. The cones may also burst open in hot weather. [13]

In its native range, P. radiata is associated with characteristic flora and fauna. It is the co-dominant canopy tree, together with Cupressus macrocarpa , which also naturally occurs only in coastal Monterey County. [14] Furthermore, one of the pine forests in Monterey, California, was the discovery site for Hickman's potentilla, an endangered species. Piperia yadonii , a rare species of orchid, is endemic to the same pine forest adjacent to Pebble Beach. In its native range, P. radiata is a principal host for the dwarf mistletoe Arceuthobium littorum . [15]

The habitat of the pines on Cedros Island contrasts greatly with the desert scrub on other parts of the island, forming zones of abrupt transition. The numerous groves form a mostly-monotypic forest of the species, with very few other plants besides seedlings emerging in the understory. In some areas, the edges of the forest form a zone that supports chaparral species, including Malosma laurina , Diplacusstellatus, and the endemic Eriogonum molle . With their large surfaces to condense fog, the pines create irrigation for themselves and their associates. Towards the far northern end of the island, a succulent community mostly consisting of Dudleya is found, and the endemic Dudleya pachyphytum can sometimes be found growing under the pines in the ecotone. [16]

A remnant P. radiata stand in Pacific Grove, the Monarch Grove Sanctuary, is a prime wintering habitat of the monarch butterfly. [17]

In South Africa, the tree is a threat to already scarce water resources. [18] The tree has remarkable roots. Monterey pine roots will reach downward as far as physically permitted by subterranean conditions. Roots have been discovered up to 12 meters (39 ft) long. [19] Efforts to remove large quantities of the non-native tree in areas of South Africa have resulted in significant increases in accessible water. [18]

Conservation status

Fungal disease

The three remaining wild stands of var. radiata (Monterey pine proper) are infected and under threat of extirpation from pine pitch canker caused by Fusarium circinatum , a fungal disease native to the southeast United States and found (in 1986) to have been introduced to California. When trees begin to die of the disease, they attract bark beetles which provide a pathway for infection of other trees. In some stands, 80–90% of trees are infected. If the disease is introduced in agroforestry areas dependent upon P. radiata, such as New Zealand, it could have catastrophic effects in those countries as well. [6]

Sphaeropsis blight (Diplodia pinea) infects P. radiata in California and causes serious damage to plantations of the species in New Zealand, Australia and South Africa, [20] especially after hail damage to growing tips.[ citation needed ]

Baja California

Pinus radiata var. binata Pinus radiata binata.jpg
Pinus radiata var. binata

On Guadalupe Island, var. binata is critically endangered. Most of the population was destroyed as tens of thousands of feral goats ate binata seedlings and caused soil erosion from the mid-19th century until just a few years ago.[ when? ] The older trees gradually died off until by 2001–2002 the population stood at only one hundred. With a program to remove the goats essentially complete by 2005, hundreds of young Guadalupe pines have started to grow up in habitat fenced after 2001, the first significant new growth in about 150 years. Possible accidental introduction of pine pitch canker is considered the biggest threat at present to the survival of the Guadalupe Island pine population. [21] The University of California's Russell Reservation forestry research station hosts an orchard planted with 73 P. radiata seedlings from Guadalupe Island and plays an important role in conserving the binata variety. [22]

Cultivation

Australia

Monterey Pine trees in Kuitpo Forest near Adelaide in South Australia Winter sun through trees - panoramio.jpg
Monterey Pine trees in Kuitpo Forest near Adelaide in South Australia
Monterey Pine trees plantation in Prospect Hill, Pemulwuy near Sydney in New South Wales Prospecthillforest.jpg
Monterey Pine trees plantation in Prospect Hill, Pemulwuy near Sydney in New South Wales

P. radiata was introduced to Australia in the 1870s. It is "the dominant tree species in the Australian plantation estate" [23] – so much so that many Australians are concerned by the resulting loss of native wildlife habitat. The species is widely regarded as an environmental weed across southeastern and southwestern Australia [24] and the removal of individual plants beyond plantations is encouraged. [25] The Kuitpo Forest, 40 kilometres (25 miles) south-east of the Adelaide city centre, is a planted forest of Monterey Pine trees. 30 kilometres (19 miles) west of the Sydney city centre, there is a forest of introduced Monterey Pine trees in Prospect Hill, in the suburb of Pemulwuy. [26]

Chile

P. radiata has greatly replaced the Valdivian temperate rain forests, where vast plantations have been planted for timber, again displacing the native forests. [27] In 2001, this species produced 5,580,724 cubic meters of lumber, or 95% of Chile's total lumber production. [28] In 2021 1.3 million of Chile's 2.3 million ha of forest plantations were planted with Pinus radiata. [29]

New Zealand

The Monterey pine (always called "Radiata Pine" or Pinus radiata in New Zealand) was first introduced into New Zealand in 1859 [30] [31] and today 89% of the country's plantation forests are of this species. [32] This includes the Kaingaroa Forest (on the central plateau of the North Island), which is one of the largest planted forests in the world. Mass plantings became common from 1900 in the Rotorua area where prison labour was used. In some areas (particularly areas that were formerly grazed that have had stock removed) it is considered an invasive species (termed a wilding conifer or more commonly wilding pine) where it has escaped from plantations. It is the most extensively used wood in New Zealand.[ citation needed ]

Use of pine in construction did not become widespread until forced by wartime shortages. It had been used in Southland from about 1920, [33] but doubts were being expressed about it as late as 1945, [34] when at least one MP considered it only suitable for interior studding. [35] Experiments in pressure treatment with water-soluble preservatives were made from 1943. [36]

Spain

In the Iberian Peninsula since the nineteenth century they have been introduced mainly in the north area in order to take advantage of their wood for the manufacture of paper pulp and for shoring work in coal mines. It is found in low altitude areas of the Autonomous Communities of Galicia, Asturias, Cantabria, the Basque Country, and in the north of the Canary Islands.[ citation needed ]

P. radiata forests have a negative effect on local ecology. In its plantations there are usually no other tree species, while its shady undergrowth does not allow the existence of a rich stratum of scrub. 13% of the wood cut annually in Spain comes from this pine. [37]

United Kingdom

The cultivar P. radiata (Aurea Group) 'Aurea' has gained the Royal Horticultural Society's Award of Garden Merit. [38] [39]

United States

P. radiata is widely used in private gardens and public landscapes in temperate California, and similar climates around the world. It is particularly commonly grown as a landscape tree in coastal areas of California outside of its native range, where the climate is virtually identical to its native range. It is fast-growing and adaptable to a broad range of soil types and climates, though it does not tolerate temperatures below about −15 °C (5 °F). Its fast growth makes it ideal for landscapes and forestry; in a good situation, P. radiata can reach its full height in 40 years or so. Though a combination of biotic and abiotic factors determines the natural distribution of P. radiata, humans have broadly expanded its distribution up and down the California coast, even reaching Oregon. [40]

Uses

Forty-five year old Pinus radiata bonsai at the National Arboretum in Canberra Bonsai pinus radiata.jpg
Forty-five year old Pinus radiata bonsai at the National Arboretum in Canberra

As timber P. radiata is suitable for a wide variety of uses, [41] and has a resinous fragrance while being worked. [42] It holds screws and nails well and takes paint and stain without difficulty, and modern kiln dried timber is very easy to work. [43] It is about 1/3 heavier than dried western red cedar. It is brittle when bent, so does not have the same load-bearing features as Oregon pine (Douglas fir, Pseudotsuga).

P. radiata is used in house construction as weatherboards, posts, beams or plywood, in fencing, retaining walls, for concrete formers. It is also used to a limited extent in boat building where untreated ply is sometimes used, but must be encased in epoxy resin to exclude moisture.

The wood is normally kiln dried to 12% moisture in 6 m (19 ft 8 in) long, clear lengths. It is available treated with a range of chemical salts, or untreated. Chemical salt treatment is well proven and such timber is frequently used in the ground as posts and poles as part of structures such as retaining walls and pole houses. The name applied to this treatment is tanalized wood. H1 and H2 treatment is suited to indoor use. H3 is the standard house timber and this grade is used for fence palings. H4 and H5 are the standard for inground use. In New Zealand a 1995 change to no longer require boron treatment in house framing timber, [44] was a key factor in the leaky homes crisis, [45] but since 2003 a series of changes have now improved the regulations. [46]

Lower grade timber is converted to pulp to make newsprint. [47] Higher grade timber is used in house construction. P. radiata is used chipped to make particle board sheets, commonly used in flooring. Other sheet products are hardboard, softboard and ply. Most ply is structural and available in 7–22 mm (0.28–0.87 in) sizes. A small amount of higher grade ply is used to produce thinner (4 and 7 mm or 0.16 and 0.28 in) ply suitable for furniture, cabinet work and boat building. This is knot and crack free and glued with resorcinol waterproof glue. Since the 1990s finger jointed joinery-grade wood has become available in up to 6 m (19 ft 8 in) lengths in a wide range of profiles.

In 1958, New Zealand boat designer Des Townson started building 186 eleven-foot (3.35 m), cold-moulded Zephyr-class dinghies, using P. radiata. In 2011 these hand-built boats fetched very high prices and were generally in excellent condition[ citation needed ].

The bark is used as a substrate for potting and re-potting orchids.

P. radiata is the most common species of Christmas tree in Australia and New Zealand.

In California, P. radiata is commonly planted to block wind or noise, or for ornamental reasons. [48]

Related Research Articles

<span class="mw-page-title-main">Pine</span> Genus of plants in the conifer family Pinaceae

A pine is any conifer tree or shrub in the genus Pinus of the family Pinaceae. Pinus is the sole genus in the subfamily Pinoideae.

<span class="mw-page-title-main">Douglas fir</span> Species of tree

The Douglas fir is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three varieties: coast Douglas-fir, Rocky Mountain Douglas-fir and Mexican Douglas-fir.

<span class="mw-page-title-main">Conifer</span> Group of cone-bearing seed plants

Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta, also known as Coniferophyta or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species.

<i>Pinus nigra</i> Species of conifer

Pinus nigra, the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as well as Crimea and in the high mountains of Northwest Africa.

<span class="mw-page-title-main">Longleaf pine</span> Species of plant (tree)

The longleaf pine is a pine species native to the Southeastern United States, found along the coastal plain from East Texas to southern Virginia, extending into northern and central Florida. In this area it is also known as "yellow pine" or "long leaf yellow pine", although it is properly just one out of a number of species termed yellow pine. It reaches a height of 30–35 m (98–115 ft) and a diameter of 0.7 m (28 in). In the past, before extensive logging, they reportedly grew to 47 m (154 ft) with a diameter of 1.2 m (47 in). The tree is a cultural symbol of the Southern United States, being the official state tree of Alabama. This particular species is one of the eight pine tree species that falls under the "Pine" designation as the state tree of North Carolina.

<i>Pinus canariensis</i> Species of conifer in the family Pinaceae

Pinus canariensis, the Canary Island pine, is a species of gymnosperm in the conifer family Pinaceae. It is a large, evergreen tree, native and endemic to the outer Canary Islands of the Atlantic Ocean.

<i>Pinus elliottii</i> Species of conifer tree

Pinus elliottii, commonly known as slash pine, is a conifer tree native to the Southeastern United States. Slash pine is named after the "slashes" – swampy ground overgrown with trees and bushes – that constitute its habitat. Other common names include swamp pine, yellow slash pine, and southern Florida pine. Slash pine has two different varieties: P. e. var. elliottii and P. e. var. densa. Historically, slash pine has been an important economic timber for naval stores, turpentine, and resin. The wood of slash pine is known for its unusually high strength, especially for a pine. It exceeds many hardwoods and is even comparable to very dense woods such as ironwood.

<i>Pinus ponderosa</i> Species of large pine tree in North America

Pinus ponderosa, commonly known as the ponderosa pine, bull pine, blackjack pine, western yellow-pine, or filipinus pine is a very large pine tree species of variable habitat native to mountainous regions of western North America. It is the most widely distributed pine species in North America.

<i>Pinus halepensis</i> Species of conifer

Pinus halepensis, commonly known as the Aleppo pine, also known as the Jerusalem pine, is a pine native to the Mediterranean region.

<i>Pinus pinaster</i> Species of plant

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<span class="mw-page-title-main">Guadalupe Island</span> Mexican island in the Pacific Ocean

Guadalupe Island is a volcanic island located 241 kilometres off the western coast of Mexico's Baja California Peninsula and about 400 km (200 nmi) southwest of the city of Ensenada in the state of Baja California, in the Pacific Ocean. The various volcanoes are extinct or dormant. In 2005 Guadalupe Island and its surrounding waters and islets were declared a biosphere reserve to restore its vegetation and to protect its population of marine mammals and birds. The island is a popular destination for great white shark cage diving. Guadalupe Island is inhabited only by scientists, military personnel operating a weather station, and a small group of seasonal fishermen. The island is mostly arid and has very little surface water.

<i>Vitex lucens</i> Species of tree

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<i>Hesperocyparis guadalupensis</i> Island endemic species of western cypress tree

Hesperocyparis guadalupensis, commonly known as Guadalupe cypress, is a species of western cypress from Guadalupe Island in the Pacific Ocean off the western coast of Mexico's Baja Peninsula. It was previously known as Cupressus guadalupensis until 2009. It is a medium sized tree with fine green to blue-green foliage. In its native habitat it depends on water from the fogs that envelop the high northern half of the island. It became an endangered species due to feral goats living on Guadalupe Island that prevented new trees from growing for more than a century. In 2005 the goats were finally removed from its island home as part of an island restoration project. New trees are growing and other plants are beginning to recover though the future of the species is not yet assured. Guadalupe cypress is closely related to the vulnerable Tecate cypress which grows on the mainland in Baja California and southern California. It is used as an ornamental tree in Mediterranean climates, particularly in Europe, but has no other significant human uses.

<span class="mw-page-title-main">Lone Pine (tree)</span>

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<span class="mw-page-title-main">Closed-cone conifer forest</span>

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Euan G. Mason is a Professor at the School of Forestry in the University of Canterbury in Christchurch, New Zealand.

<span class="mw-page-title-main">Wilding conifer</span> Invasive trees in New Zealand

Wilding conifers, also known as wilding pines, are invasive trees in the high country of New Zealand. Millions of dollars are spent on controlling their spread.

<span class="mw-page-title-main">Forestry in New Zealand</span>

Forestry in New Zealand has a history starting with European settlement in the 19th century and is now an industry worth seven percent of annual revenue. Much of the original native forest cover was burnt off and logged, however forests have been extensively planted, predominantly with fast-growing cultivars of the Monterey Pine. Wood chips, whole logs, lumber and paper products are exported from New Zealand.

<i>Fusarium circinatum</i> Species of fungus

Fusarium circinatum is a fungal plant pathogen that causes the serious disease pitch canker on pine trees and Douglas firs. The most common hosts of the pathogen include slash pine, loblolly pine, Monterey pine, Mexican weeping pine, and Douglas fir. Like other Fusarium species in the phylum Ascomycota, it is the asexual reproductive state of the fungus and has a teleomorph, Gibberella circinata.

<i>Hylastes ater</i> Species of beetle

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