Parasaurus

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Parasaurus
Temporal range: Wuchiapingian, 258.9–255.7  Ma
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Parasaurus NT.png
Restoration
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Parareptilia
Order: Procolophonomorpha
Clade: Pareiasauria
Family: Pareiasauridae
Genus: Parasaurus
von Meyer, 1857
Type species
Parasaurus geinitzi
von Meyer, 1857

Parasaurus (meaning "near lizard") is a genus of pareiasaur known from fossils collected in the Kupferschiefer in Germany (Hesse, Thuringia and Lower Saxony), dating to the Late Permian (Wuchiapingian). The type species, Parasaurus geinitzi, described by Hermann von Meyer in 1857, [1] was the first pareiasaur ever described. [2] The seven known specimens were redescribed in 2008. [2]

Contents

Discovery and naming

As early as 1848, pareiasaur fossils have been reported from Germany; [3] some of these fossils may have belonged to Parasaurus. [4] It was not until 1857 when von Meyer described these fossils and created the Parasaurus genus. [1] von Meyer classified Parasaurus as a reptile but it was classified as a pareiasaur when the family was created in 1888. [5] [4] Lee (1997) classified Parasaurus as a nomen dubium . In 2008, Tsuji and Müller re-evaluated the genus. [2]

Description

Parasaurus was small for a pareiasaur, only around 50 centimetres (1.6 ft) long. Axial osteoderms appear to be absent. The skull surface is pitted, with small spike-like horns on the supratemporal and quadratojugal. [2]

Phylogeny

Below is a cladogram from Tsuji et al. (2013): [6]

Pareiasauria

"Bradysaurus" seeleyi

Bradysaurus baini

Nochelesaurus

Embrithosaurus

Bunostegos

Deltavjatia

Parasaurus

Velosauria

Nanopareia

Provelosaurus

Anthodon

Pumiliopareia

Shansisaurus

Shihtienfenia

Pareiasuchus peringueyi

Pareiasuchus nasicornis

Arganaceras

Elginia

Obirkovia

Pareiasaurus

Sanchuansaurus

Scutosaurus

Paleoenvironment

The Kupferschiefer is a marine unit that forms part of the Zechstein, a sequence of rocks formed on the edge of the Zechstein Sea, a large inland shallow sea that existed in Northern Europe during the Late Permian. The environment at the time of deposition is considered to have been semi-arid. The terrestrial flora of the Zechstein is dominated by conifers, with seed ferns also being common, while taeniopterids, ginkgophytes and sphenophytes are rare. Other terrestrial vertebrates found in the Kupferschiefer and lower Zechstein include the gliding weigeltisaurid reptiles Weigeltisaurus and Glaurung, the archosauromorph reptile Protorosaurus, the cynodont Procynosuchus, and indeterminate captorhinids, dicynodonts, and dissorophid temnospondyls. [7] [8] [9] [10]

Related Research Articles

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Pareiasaurs are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with osteoderms which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event.

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<i>Coelurosauravus</i> Extinct genus of reptiles

Coelurosauravus is an extinct genus of gliding reptile, known from the Late Permian of Madagascar. Like other members of the family Weigeltisauridae, members of this genus possessed long, rod-like ossifications projecting outwards from the body. These bony rods were not extensions of the ribs but were instead a feature unique to weigeltisaurids. It is believed that during life, these structures formed folding wings used for gliding flight, similar to living gliding Draco lizards.

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<i>Weigeltisaurus</i> Extinct genus of reptiles

Weigeltisaurus is an extinct genus of weigeltisaurid reptile from the Late Permian Kupferschiefer of Germany and Marl Slate of England. It has a single species, originally named as Palaechamaeleo jaekeli in 1930 and later assigned the name Weigeltisaurus jaekeli in 1939, when it was revealed that Palaeochamaeleo was a preoccupied name. A 1987 review by Evans and Haubold later lumped Weigeltisaurus jaekeli under Coelurosauravus as a second species of that genus. A 2015 reassessment of skull morphology study substantiated the validity of Weigeltisaurus and subsequent authors have used this genus. Like other Weigeltisaurids, they possessed long rod-like bones that radiated from the trunk that were likely used to support membranes used for gliding, similar to extant Draco lizards.

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References

  1. 1 2 Meyer, H.V. (1857). Beiträge zur näheren Kenntnis fossiler Reptilien. Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde, 1857: 103–104.
  2. 1 2 3 4 L. A. Tsuji and J. Müller. (2008). A Re-evaluation of Parasaurus geinitzi, the first named pareiasaur (Amniota, Parareptilia). Canadian Journal of Earth Sciences 45(10):1111-1121
  3. Geinitz, H.B. (1848). Die Versteinerungen des deutschen Zechsteingebirges. In Die Verteinerungen des Zechsteingebirges und Rothliegenden oder des permischen Systemes in Sachsen. Edited by H.B. Geinitz and A. von Gutbier. Arnoldische Buchhandlung, Dresden and Leipzig, pp. 1–29. Google Scholar
  4. 1 2 Society of Vertebrate Paleontology (24 June 2013). "Pareiasaur: Bumpy beast was a desert dweller". ScienceDaily. Retrieved 17 May 2014.
  5. H. G. Seeley. (1888). Croonian Lecture: Researches on the Structure, Organization, and Classification of the Fossil Reptilia. II. On Pareiasaurus bombidens (Owen), and the Significance of Its Affinities to Amphibians, Reptiles, and Mammals. Philosophical Transactions of the Royal Society of London B 179:59-109
  6. Tsuji, L. A.; Sidor, C. A.; Steyer, J. - S. B.; Smith, R. M. H.; Tabor, N. J.; Ide, O. (2013). "The vertebrate fauna of the Upper Permian of Niger—VII. Cranial anatomy and relationships of Bunostegos akokanensis (Pareiasauria)". Journal of Vertebrate Paleontology. 33 (4): 747–763. doi:10.1080/02724634.2013.739537. S2CID   86097405.
  7. Bernardi, Massimo; Petti, Fabio Massimo; Kustatscher, Evelyn; Franz, Matthias; Hartkopf-Fröder, Christoph; Labandeira, Conrad C.; Wappler, Torsten; Van Konijnenburg-Van Cittert, Johanna H.A.; Peecook, Brandon R.; Angielczyk, Kenneth D. (2017-12-01). "Late Permian (Lopingian) terrestrial ecosystems: A global comparison with new data from the low-latitude Bletterbach Biota". Earth-Science Reviews. 175: 18–43. Bibcode:2017ESRv..175...18B. doi: 10.1016/j.earscirev.2017.10.002 . ISSN   0012-8252. S2CID   134260553.
  8. Witzmann, Florian (2005-05-31). "A dissorophid temnospondyl in the Upper Permian Kupferschiefer of Germany". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 2005 (5): 289–300. doi:10.1127/njgpm/2005/2005/289. ISSN   0028-3630.
  9. Sues, Hans-Dieter; Munk, Wolfgang (March 1996). "A remarkable assemblage of terrestrial tetrapods from the Zechstein (Upper Permian: Tatarian) near Korbach (northwestern Hesse)". Paläontologische Zeitschrift. 70 (1–2): 213–223. doi:10.1007/BF02988279. ISSN   0031-0220. S2CID   128465809.
  10. Bulanov, V. V.; Sennikov, A. G. (2015-12-01). "Glaurung schneideri gen. et sp. nov., a new weigeltisaurid (Reptilia) from the Kupferschiefer (Upper Permian) of Germany". Paleontological Journal. 49 (12): 1353–1364. doi:10.1134/S0031030115120035. ISSN   1555-6174. S2CID   87461613.