Interior Alaskan wolf

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Interior Alaskan wolf
Wolf on the move.jpg
Interior Alaskan wolf in Denali National Park
Status TNC G4.svg
Apparently Secure  (NatureServe) [1]
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Canidae
Genus: Canis
Species:
Subspecies:
C. l. pambasileus
Trinomial name
Canis lupus pambasileus
Elliot, 1905 [2]
North American gray wolf subspecies distribution according to Goldman (1944) & MSW3 (2005).png
Historical and present range of gray wolf subspecies in North America

The Interior Alaskan wolf [3] [4] (Canis lupus pambasileus), also known as the Yukon wolf, is a subspecies of gray wolf native to parts of British Columbia, the Northwest Territories, [5] Interior Alaska and Yukon. [4]

Contents

Taxonomy

It is recognized as a subspecies of Canis lupus in the taxonomic authority Mammal Species of the World (2005). [6] It was first described in 1905 by the American zoologist Daniel Elliot as Canis pambasileus and with the name "autocrat timber wolf", based on a specimen from the Susitna River, Mount McKinley region, Alaska. Elliot distinguishes this wolf by the teeth in both jaws being large and heavy, and along with the skull exceed those of C. l. occidentalis (the Northwestern wolf) of a comparable body size. This wolf's coat ranges from black to white or a mix of both. [2] In 1944, the American zoologist Edward Goldman recognized this wolf as Canis lupus pambasileus Elliot, 1905 and with the name "Interior Alaskan wolf". [3]

Physiology

Description

The wolf has a height of 85 cm (33.5 in)[ citation needed ] and a length between 150 and 196 cm (59 and 77 in)[ citation needed ] with an average male weight of 56.3 kilograms (124 lb) and for females 38.5 kilograms (85 lb). Individual weights can vary from 32 kilograms (71 lb) to 60 kilograms (130 lb). [5] In Yukon-Charley Rivers National Preserve male wolves average 56.3 kg (111 lb) and females 44 kg (97 lb); in Denali National Park and Preserve male wolves average 52.6 kg (105 lb). [7] One specimen weighed 81.4 kilograms (179 lb). [8] [9] It was killed on 70 Mile River in east-central Alaska on July 12, 1939. [10] One wolf was purported to weigh 212 pounds (96 kg), however large Alaskan wolves are known to hold up to 20 pounds (9.1 kg) of moose meat in their stomachs. [11] The most common color for this subspecies is tawny grey or tan, but can also range from white to black. The lifespan ranges from 4 to 10 years, the oldest being 12 years. [5]

Pack habits

Average pack size is 7 to 9 wolves, but can vary; like other wolves, the pack consists of a mated pair and their offspring. The pair is usually the only ones that breed. A wolf that has left its pack may travel up to 500 km (310.7 mi) to breed. The minimum breeding age is 1 year, and the average litter size is 4–6 pups. [12]

Disease vectors

While exact data is not available, it is suspected that various types of diseases, such as rabies and distemper, affect this subspecies, sometimes to the point that the stability of the subspecies is changed in parts of its region. [13]

Distribution and habitat

This wolf is native to the interior of Alaska, United States, and the Yukon, Canada save for the tundra region of the Arctic Coast. [4] Yukon wolves' main habitats are boreal forests, alpine, subalpine, and Arctic tundra. The population in Canadian Yukon is estimated to be 5,000, which ranges in all of Yukon except for Kluane National Park. The population density depends on prey, with the densest population being in Teslin, Yukon, Canada where there are 9 wolves per 1,000 square kilometers (386.1 sq mi), while the least dense is in Northern Yukon, with 3 wolves per 1,000 square kilometers. [12]

Diet

Feeding on caribou in Denali National Park and Preserve NPS Photo Cindy Gonterman (5752389453).jpg
Feeding on caribou in Denali National Park and Preserve

The diet of this wolf varies by region – moose is the main prey in southern Yukon, followed by boreal woodland caribou and Dall sheep. Barren-ground caribou is main prey in the North Slope. When hunting moose, wolves mainly kill calves and old moose when fleeing. Usually the hunting success rate for moose is 10% in one hunt, and a pack usually kill a moose every 5 to 6 days to eat for 2 – 3 days. Moose are more likely to stand their ground than caribou, which tend to flee, decreasing their survival rate. Wolves usually kill a caribou every 3 days during winter and eat for a day. Dall sheep are common prey in Kluane Game Sanctuary and National Park when moose and caribou are not available. [12]

History

In pre-colonial Canada, the local Aboriginal population hunted this wolf for its fur. This continued into the 1800s, with colonists selling wolf furs to Aboriginal tribes in the area, who used them to line their clothing. [14]

The first true mapping of the wolf population in the Yukon began in the 1950s and, subsequently, a program of wolf-poisoning began. This came about because of the public stigma regarding wolves during the time. [15]

Yukon Conservation and Management Plan

A plan known as the Yukon Wolf Conservation and Management Plan was created in the 1980s by the Yukon government in order to determine a method to control the population of wolves in Yukon. The studies that would be conducted wherein a reduction in the wolf population would only occur in one specified testing area at a time. Also no hunting would be allowed within the testing area, in order to have a strict control and understanding of the results of any study. The studies would also be conducted in only 7 of the 23 ecoregions in Yukon. [16]

The aftereffects of the plan in the 2000s involved continued study and the creation of "rigid guidelines for ensuring long-term wolf conservation" and limited the amount of wolf control enacted by decreasing the scope and length of any future studies. The planning group also set forth methods of increasing public awareness of the importance of wolves and making more strict wolf hunting laws. The plan was eventually opposed by environmental groups who did not wish for wolf control in any form. [17] [18]

Finlayson Recovery Program (1983–1992)

The first control program was started because of observations and complaints by the people of Ross River that the Filayson herd of caribou has been declining in size. In response, the government, from the beginning of the program to 1989, reduced the number of wolf packs in the area from 25 to 7, which involved reducing the number of individual members from 215 to 29. In order to test if a concurrent effort would have a greater effect, hunting any caribou in the herd was also limited and controlled by a strict permit system. [19]

The result was that the number of caribou more than doubled and the number of wolves returned to their previous count before the control period. Since the end of the program, the number of caribou has begun to steadily decline, though the health of the herd remained stable regardless. While the government had wished to continue efforts at wolf control in the area, an agreement could not be made with the First Nation peoples of Ross River, so the effort was eventually abandoned. [19]

Aishihik Recovery Program (1992–1997)

A five-year control program created as a part of the Yukon Wolf Plan ended in 1997 that detailed the extermination of around 80% of the Yukon wolf population within southwest Yukon, specifically around the Aishihik area. This was accomplished by aerial wolf hunting and through the use of snares [15] and traps. Numerous wolves were also neutered in the years between 1994 and 1997. After recovering the bodies of slain wolves, the pelts were removed and sold. [16] [20]

The program was a test on whether decreased wolf population would cause the population of caribou, moose, and Dall sheep to increase. [16] [21] Subsequently, it could be decided that the major limiting factor in terms of the population growth of these species is attributed to predation by the Yukon wolf. The resulting evidence showed that, while the Yukon wolf greatly affected and controlled the population of moose, the survival rate of adult caribou was not affected by the decrease in the number of wolves in the area, nor was the Dall sheep population affected at all. Thus, the study concluded that the Yukon wolf, along with hunting by humans, was one of the greatest controlling factors for the moose population and for the caribou population as a whole. [16]

The study also concluded that, in future testing areas, lethal methods of reducing wolf populations would be minimized, as it was clearly found during the study that sterilization was far more effective in controlling the population and it also did not affect wolf behavior. [16]

In 1994, during the study, a big game outfitter was convicted in court of attempting to convince guides in the area to place poison on the ground. Some poisoned Yukon wolves and birds were found in the Aishihik testing area and 4 of the packs in the area lowered from 38 individual members to 8 members. [16]

Chisana Recovery Program (1996–present)

The third study and control program under the Yukon Wolf Plan began under the insistence of the White River and the Kluane First Nation in regards to the Chisana caribou herd that roamed around the Saint Elias Mountains in the area. They had observed that caribou numbers has dropped from over 3000 in the early 1960s to around 400 by the 1990s. The Yukon government's first response was to inactivate all hunting permits in the area. Afterwards, various studies were conducted on the herd in order to determine what could be causing it to continuously decline. [19]

Both the government and the local community were against enacting another aerial wolf hunting program and, so, the Yukon wolves were left alone in the area. Instead, a process of helping breed caribou was enacted that was fairly successful. Further breeding attempts and studies are still ongoing. [19]

2009 aerial wolf hunt

In early 2009, the US Board of Game passed through a plan by the Alaska Department of Fish and Game that allowed " the state to hire private helicopters to kill wolves." The plan itself had been given to the Board of Game after the period of public comment has passed, which made it so that the public had no say in the plan or its workings. The Alaska division of the Defenders of Wildlife was outraged over these actions, stating that "the Palin administration and the Board of Game are acting with complete disregard for scientific wildlife management." [22]

A few days later, a government-sanctioned aerial wolf hunt began in Alaska. It was initiated within only a few hours after the Alaska Department of Fish and Game told the National Park Service that such an effort was going to be undertaken in the area around Tanana. With about 200 wolves being the goal stated for the hunt, more than 30 were killed within the first day, including those that had been radio-collared for research purposes. The National Park Service was greatly displeased with the loss of collared wolves, as they had been part of a 20-year study. [22]

Before the hunt began, the National Park Service released a statement that said that, if the hunting plan was continued to completion, "this would leave one-to-two wolves per 1,000 square kilometers in the Upper Yukon Wolf Control Area, approximating the lowest known wolf population densities in Alaska." [22]

Related Research Articles

<span class="mw-page-title-main">Wolf</span> Type of canine

The wolf, also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of Canis lupus have been recognized, including the dog and dingo, though gray wolves, as popularly understood, only comprise naturally-occurring wild subspecies. The wolf is the largest extant member of the family Canidae, and is further distinguished from other Canis species by its less pointed ears and muzzle, as well as a shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller Canis species, such as the coyote and the golden jackal, to produce fertile hybrids with them. The wolf's fur is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white.

<span class="mw-page-title-main">Kluane National Park and Reserve</span> National park and park reserve in Yukon, Canada

Kluane National Park and Reserve are two protected areas in the southwest corner of the territory of Yukon. The National Park Reserve was set aside in 1972 to become a national park, pending settlement of First Nations land claims. It covered an area of 22,013 square kilometres. When agreement was reached with the Champagne and Aishihik First Nations over an eastern portion of the Reserve, that part—about 5,900 square kilometres —became a national park in 1993, and is a unit of the national park system administered co-operatively with Parks Canada. The larger western section remains a Reserve, awaiting a final land claim settlement with the Kluane First Nation. The park borders British Columbia to the south, while the Reserve borders both British Columbia to the south, and the United States (Alaska) to the south and west.

<span class="mw-page-title-main">Eastern wolf</span> Subspecies of carnivore

The eastern wolf, also known as the timber wolf, Algonquin wolf and eastern timber wolf, is a canine of debated taxonomy native to the Great Lakes region and southeastern Canada. It is considered to be either a unique subspecies of gray wolf or red wolf or a separate species from both. Many studies have found the eastern wolf to be the product of ancient and recent genetic admixture between the gray wolf and the coyote, while other studies have found some or all populations of the eastern wolf, as well as coyotes, originally separated from a common ancestor with the wolf over 1 million years ago and that these populations of the eastern wolf may be the same species as or a closely related species to the red wolf of the Southeastern United States. Regardless of its status, it is regarded as unique and therefore worthy of conservation with Canada citing the population in eastern Canada as being the eastern wolf population subject to protection.

<span class="mw-page-title-main">Dall sheep</span> Species of mammal

Ovis dalli, also known as the Dall sheep or thinhorn sheep, is a species of wild sheep native to northwestern North America. Ovis dalli contains two subspecies: Ovis dalli dalli and Ovis dalli stonei. O. dalli live in mountainous alpine habitats distributed across northwestern British Columbia, the Yukon, Northwest Territories and Alaska. They browse a variety of plants such as grasses, sedges and even shrubs such as willow, during different times of the year. They also acquire minerals to supplement their diet from mineral licks. Like other Ovis species, the rams engage in dominance contests with their horns.

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<span class="mw-page-title-main">Hunting behavior of gray wolves</span>

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References

  1. "Grey Wolf" . Retrieved March 16, 2015.
  2. 1 2 Descriptions of Three Apparently New Species of Mammals Elliot, Proc. Biol. Sci. Wash., XVIII 1905, p79
  3. 1 2 Young, Stanley P.; Goldman, Edward A. (1944). The Wolves of North America. Vol. 2. Dover Publications, New York. pp. 413–477. ISBN   978-0486211930.
  4. 1 2 3 Mech, L. David (1981), The Wolf: The Ecology and Behaviour of an Endangered Species, University of Minnesota Press, pp. 352–353, ISBN   0-8166-1026-6
  5. 1 2 3 "Gray wolf (in the Yukon)" (PDF). Environment Yukon. Government of Canada. 2017. Retrieved 17 April 2017.
  6. Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 575–577. ISBN   978-0-8018-8221-0. OCLC   62265494. url=https://books.google.com/books?id=JgAMbNSt8ikC&pg=PA576
  7. "Big Wolves and Ordinary Wolves, Alaska Department of Fish and Game".
  8. The Wolves of Isle Royale. Fauna Series 7. Fauna of the National Parks of the United States. 1966. p. 76. ISBN   1-4102-0249-6.
  9. Young, Stanley P.; Goldman, Edward A. (1944). The Wolves of North America, Part II. New York, Dover Publications, Inc.
  10. Lopez, Barry H. (1978). Of Wolves and Men. J. M. Dent and Sons Limited. p. 18. ISBN   0-7432-4936-4.
  11. Jim Rearden (2002). The Wolves of Alaska: A Fact-based Saga. Portland Oregon: Alaska Northwest Books. p. 131. ISBN   978-0-88240-933-7.
  12. 1 2 3 Environment Yukon – Grey wolf
  13. Ludwig N. Carbyn (1983). Wolves in Canada and Alaska: their status, biology, and management : proceedings of the Wolf Symposium held in Edmonton, Alberta, 12–14 May 1981. Volume 45 of Canadian wildlife service report series, Canadian Wildlife Service Ottawa. Canadian Wildlife Service. ISBN   9780660113784.
  14. William Watson Woollen; Paul Leland Haworth (1924). Paul Leland Haworth (ed.). The inside passage to Alaska, 1792–1920: with an account of the North Pacific coast from Cape Mendocino to Cook Inlet, from the accounts left by Vancouver and other early explorers, and from the author's journals of exploration and travel in that region, Volume 2. The Arthur H. Clark Co.
  15. 1 2 Bob Jickling (2005). ""The wolf must not be made a fool of": reflections on education, ethics, and epistemology". In Peggy Tripp; Linda June Muzzin (eds.). Teaching as activism: equity meets environmentalism. McGill-Queen's Press. pp.  35–45. ISBN   978-0-7735-2808-6.
  16. 1 2 3 4 5 6 Robert D. Hayes, Richard Farnell, Richard M. P. Ward, Jean Carey, Michael Dehn, Gerald W. Kuzyk, Alan M. Baer, Craig L. Gardner & Mark O'Donoghue (2003). "Experimental reduction of wolves in the Yukon: ungulate responses and management implications". Wildlife Monographs . 152 (152): 1–35. JSTOR   3830836.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  17. S. H. Fritts; R. O. Stephenson; R. D. Hayes & L. Boitani (2003). "Wolves and humans". In L. David Mech & Luigi Boitani (eds.). Wolves: Behavior, Ecology, and Conservation. University of Chicago Press. pp. 289–316. ISBN   978-0-226-51696-7.
  18. "Yukon wolf kill sparks outrage". Chicago Tribune. December 21, 1994. Retrieved August 28, 2010.[ permanent dead link ]
  19. 1 2 3 4 Kelly Hayes & Gerry Couture (March 2–6, 2004). Community Involvement in Recovering Woodland Caribou Populations: Yukon Success Stories (PDF). Species at Risk 2004 Pathways to Recovery Conference. Victoria, British Columbia.
  20. Bob Jickling & Paul C. Paquet (2005). "Wolf stories: Reflections on science, ethics, and epistemology". Environmental Ethics . 27 (2): 115–134. doi:10.5840/enviroethics200527226.
  21. Todd Wilkinson (August 1994). "Bison flapping in the breeze". Backpacker . 22 (134): 18.
  22. 1 2 3 Defenders of Wildlife (March 16, 2009). "State Initiates Massive New Helicopter Wolf Slaughter". Common Dreams.