Helveticosauridae

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Helveticosauridae
Temporal range: Middle Triassic, 243–241  Ma
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Eusaurosphargis dalsassoi.jpg
Eusaurosphargis dalsassoi PIMUZ A/III 4380
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauromorpha (?)
Family: Helveticosauridae
Peyer, 1955
Genera

Helveticosauridae is an extinct family of basal marine reptiles known from the Middle Triassic (Anisian-Ladinian boundary) of southern Switzerland and northern Italy. [1] [2]

The type species of the family is Helveticosaurus zollingeri , named by Bernhard Peyer in 1955 based on a single nearly complete specimen T 4352 collected at Cava Tre Fontane from the Anisian-Ladinian boundary of Monte San Giorgio, Switzerland. Peyer (1955) considered the species to be a very distinctive member of the order Placodontia, and thus erected Helveticosauridae as well as the superfamily Helveticosauroidea to contain it within Placodontia. [1]

Nosotti and Rieppel (2003) described Eusaurosphargis from the equivalent beds at Cava di Besano of the Besano Formation (Anisian-Ladinian boundary) of Italy. Their phylogenetic analysis recovered it as the sister taxon of Helveticosaurus, and thus it was assigned to Helveticosauridae. Based on the description in the literature available for Saurosphargis (whose holotype is lost), they considered it to also fall in the Helveticosauridae clade. [2] The anatomy of Saurosphargis was finally clarified by comparisons with the well-preserved specimens of Sinosaurosphargis , and as a result Saurosphargis was no longer considered to be a nomen dubium , and thus could be included in a phylogenetic analysis. Li et al. (2011) found Saurosphargis and Sinosaurosphargis to form a clade separate from that of Eusaurosphargis and Helveticosaurus, thus Saurosphargis was removed from Helveticosauridae, and placed in its own family Saurosphargidae together with Sinosaurosphargis. [3]

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Eusaurosphargis is an extinct genus of a diapsid reptile, known from the Middle Triassic Besano Formation of northern Italy and Prosanto Formation of south-eastern Switzerland. It contains a single species, Eusaurosphargis dalsassoi. It was a small reptile, measuring 20 cm (7.9 in) long.

The Besano Formation is a geological formation in the southern Alps of northwestern Italy and southern Switzerland. This formation, a thin but fossiliferous succession of dolomite and black shale, is famous for its preservation of Middle Triassic (Anisian–Ladinian) marine life including fish and aquatic reptiles. It is exposed in the Monte San Giorgio and Besano area. It is among the formations responsible for the area being designated as a UNESCO World Heritage Site. In Switzerland, it is also known as the Grenzbitumenzone. The Anisian-Ladinian boundary lies in the upper part of the Besano Formation.

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References

  1. 1 2 Bernhard Peyer (1955). "Die Triasfauna der Tessiner Kalkalpen. XVIII. Helveticosaurus zollingeri, n.g. n.sp". Schweizerische Paläontologische Abhandlungen. 72: 3–50.
  2. 1 2 Nosotti, Stefania; Rieppel, Olivier (2003). "Eusaurosphargis dalsassoi n. gen. n. sp., a new, unusual diapsid reptile from the Middle Triassic of Besano (Lombardy, N Italy)". Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano Ile. 31 (3): 1–33.
  3. Li, Chun; Olivier Rieppel; Xiao-Chun Wu; Li-Jun Zhao; Li-Ting Wang (2011). "A new Triassic marine reptile from southwestern China". Journal of Vertebrate Paleontology. 31 (2): 303–312. doi:10.1080/02724634.2011.550368.