The Triassic is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.
Lepidosauromorpha is a group of reptiles comprising all diapsids closer to lizards than to archosaurs. The only living sub-group is the Lepidosauria, which contains two subdivisions, Squamata, which contains lizards and snakes, and Rhynchocephalia, the only extant species of which is the tuatara.
A number of animals are capable of aerial locomotion, either by powered flight or by gliding. This trait has appeared by evolution many times, without any single common ancestor. Flight has evolved at least four times in separate animals: insects, pterosaurs, birds, and bats. Gliding has evolved on many more occasions. Usually the development is to aid canopy animals in getting from tree to tree, although there are other possibilities. Gliding, in particular, has evolved among rainforest animals, especially in the rainforests in Asia where the trees are tall and widely spaced. Several species of aquatic animals, and a few amphibians and reptiles have also evolved this gliding flight ability, typically as a means of evading predators.
Coelurosauravus is an extinct genus of gliding reptile, known from the Late Permian of Madagascar. Like other members of the family Weigeltisauridae, members of this genus possessed long, rod-like ossifications projecting outwards from the body. These bony rods were not extensions of the ribs but were instead a feature unique to weigeltisaurids. It is believed that during life, these structures formed folding wings used for gliding flight, similar to living gliding Draco lizards.
Avicephala is a potentially polyphyletic grouping of extinct diapsid reptiles that lived during the Late Permian and Triassic periods characterised by superficially bird-like skulls and arboreal lifestyles. As a clade, Avicephala is defined as including the gliding weigeltisaurids and the arboreal drepanosaurs to the exclusion of other major diapsid groups. This relationship is not recovered in the majority of phylogenetic analyses of early diapsids and so Avicephala is typically regarded as an unnatural grouping. However, the clade was recovered again in 2021 in a redescription of Weigeltisaurus, raising the possibility that the clade may be valid after all.
Icarosaurus is an extinct genus of kuehneosaurid reptile from the Late Triassic Lower Lockatong Formation of New Jersey. It is closely related to lizards and the tuatara. Based on a partial skeleton missing part of the tail, some ribs, a hand, and parts of the legs, it was a small animal, about 10 centimeters (4 in) long from the skull to the hips. Like its relative Kuehneosaurus, it was able to glide short distances using 'wings' consisting of highly elongated ribs covered with skin. These gliding membranes would have had a convex upper surface and a concave lower surface, thus creating a simple airfoil structure well-suited to gliding. This method of gliding is also seen in Coelurosauravus and the modern Draco, neither of which are closely related to Icarosaurus.
Weigeltisaurus is an extinct genus of weigeltisaurid reptile from the Late Permian Kupferschiefer of Germany and Marl Slate of England. It has a single species, originally named as Palaechamaeleo jaekeli in 1930 and later assigned the name Weigeltisaurus jaekeli in 1939, when it was revealed that Palaeochamaeleo was a preoccupied name. A 1987 review by Evans and Haubold later lumped Weigeltisaurus jaekeli under Coelurosauravus as a second species of that genus. A 2015 reassessment of skull morphology study substantiated the validity of Weigeltisaurus and subsequent authors have used this genus. Like other Weigeltisaurids, they possessed long rod-like bones that radiated from the trunk that were likely used to support membranes used for gliding, similar to extant Draco lizards.
Megalancosaurus is a genus of extinct reptile from the Late Triassic Dolomia di Forni Formation and Zorzino Limestone of northern Italy, and one of the best known drepanosaurids. The type species is M. preonensis; a translation of the animal's scientific name would be "long armed reptile from the Preone Valley."
Drepanosaurs are a group of extinct reptiles that lived between the Carnian and Rhaetian stages of the late Triassic Period, approximately between 230 and 210 million years ago. The various species of drepanosaurid were characterized by specialized grasping limbs and often prehensile tails, adaptions for arboreal (tree-dwelling) and fossorial (digging) lifestyles, with some having also been suggested to be aquatic. Fossils of drepanosaurs have been found in Arizona, New Mexico, New Jersey, Utah, England, and northern Italy. The name is taken from the family's namesake genus Drepanosaurus, which means "sickle lizard," a reference to their strongly curved claws.
Xianglong is a genus of Cretaceous lizard discovered in the Zhuanchengzi, near Yizhou, Yixian, Liaoning Province of China. It is known from LPM 000666, a single complete skeleton with skin impressions. The specimen comes from the Barremian-age Lower Cretaceous Yixian Formation, near Yizhou. The most notable feature about Xianglong is its bizarre oversized ribs, eight on each side, which were attached to a membrane of body tissue and allowed the lizard to glide. While in its original description it was considered to acrodont lizard, with a cladistic analysis in the same study suggesting that it was grouped with iguanians such as agamines, chamaeleonids, and leiolepidines, it was later shown that this was due to misinterpretation of the crushed skull, and its affinities with other lizards remains uncertain.
Mecistotrachelos is an extinct genus of gliding reptile from the Late Triassic of Virginia. It is generally interpreted as an archosauromorph, distantly related to crocodylians and dinosaurs. The type and only known species is M. apeoros. This specific name translates to "soaring longest neck", in reference to its gliding habits and long neck. This superficially lizard-like animal was able to spread its lengthened ribs and glide on wing-like membranes. Mecistotrachelos had a much longer neck than other gliding reptiles of the Triassic such as Icarosaurus and Kuehneosaurus. It was probably an arboreal insectivore.
Eolacertilia is an extinct clade of lepidosauriform diapsid reptiles known from the Late Permian to the Late Triassic. It is uncertain as to whether they are a natural group and it has been suggested that they form a "waste basket" taxon. Currently, the only members of the group are Paliguana and Kuehneosauridae. Other genera were transferred to basal groups within Diapsida, Archosauromorpha.
Kuehneosauridae is an extinct family of small, lizard-like gliding diapsids known from the Triassic period of Europe and North America.
Weigeltisauridae is a family of gliding neodiapsid reptiles that lived during the Late Permian, between 259.51 and 251.9 million years ago. Fossils of weigeltisaurids have been found in Madagascar, Germany, Great Britain, and Russia. They are characterized by long, hollow rod-shaped bones extending from the torso that probably supported wing-like membranes. Similar membranes are also found in several other extinct reptiles such as kuehneosaurids and Mecistotrachelos, as well as living gliding lizards, although each group evolved these structures independently.
Pamelina is an extinct genus of basal kuehneosaurid known from Early Triassic rocks of Czatkowice 1, Poland. It was first named by Susan E. Evans in 2009 and the type species is Pamelina polonica. It is the oldest known member of Kuehneosauridae. The vertebrae have characteristics consistent with gliding or parachuting.
Kuehneosuchus is an extinct genus of Late Triassic kuehneosaurid reptile known from the Late Triassic of southwest England. It was named by P. L. Robinson in 1967 and the type and only species is Kuehneosuchus latissimus. It is known from the holotype NHMUK PV R 6111, a set of associated vertebrae and ribs. It is a derived kuehneosaurid, most closely related to Kuehneosaurus. The genera are very similar and can be distinguished from one another primarily on the length of their "wing" ribs, relatively short and massive in Kuehneosaurus but up to 4 times longer and more gracile in Kuehneosuchus. However, the skull and major postcranial bones are identical in both taxa, as their age and horizon. According to aerodynamic studies Kuehneosuchus, unlike Kuehneosaurus which may be a species of the same genus or represent a different sexual morph, was probably a glider.
Azendohsauridae is a family of allokotosaurian archosauromorphs that lived during the Middle to Late Triassic period, around 242-216 million years ago. The family was originally named solely for the eponymous Azendohsaurus, marking out its distinctiveness from other allokotosaurs, but as of 2022 the family now includes four other genera: the basal genus Pamelaria, the large horned herbivore Shringasaurus, and two carnivorous genera grouped into the subfamily-level subclade Malerisaurinae, Malerisaurus and Puercosuchus, and potentially also the dubious genus Otischalkia. Most fossils of azendohsaurids have a Gondwanan distribution, with multiple species known across Morocco and Madagascar in Africa as well as India, although fossils of malerisaurine azendohsaurids have also been found in the southwestern United States of North America.
Ozimek is a genus of sharovipterygid archosauromorph reptile, known from Late Triassic deposits in Poland and closely related to the Kyrgyzstani Sharovipteryx. It contains one species, O. volans, named in 2016 by Jerzy Dzik and Tomasz Sulej. Like Sharovipteryx, Ozimek had long, slender limbs with the hindlimbs longer than the forelimbs; the hindlimbs likely supported gliding membranes as fossilized in Sharovipteryx. Another unusual characteristic was the shoulder girdle, where the massive coracoids formed a shield-like structure covering the bottom of the shoulder region that would have limited mobility. In other respects, such as its long neck, it was a typical member of the non-natural grouping Protorosauria. Phylogenetic analysis has indicated that it, possibly along with Sharovipteryx, may have been an unusual member of the protorosaur group Tanystropheidae, although further study of its anatomy is needed to resolve its precise relationships.
Pamela Lamplugh Robinson was a British paleontologist who worked extensively on the fauna of the Triassic and Early Jurassic of Gloucestershire and later worked in India on the Mesozoic and Gondwanan fauna. She helped establish the geology unit at the Indian Statistical Institute and directed research in vertebrate paleontology of India in the 1960s.
Fraxinisaura is an extinct genus of basal lepidosauromorph reptile known from the Middle Triassic of Germany. The only known species is Fraxinisaura rozynekae. It possessed an elongated snout, unique features of the teeth, and an ilium which was intermediate in orientation between sphenodontians and squamates. Based on characteristics of the maxilla, it is considered a close relative of Marmoretta from the Middle Jurassic of the United Kingdom, resolving a ghost lineage between that genus and other Triassic basal lepidosauromorphs.
